Vasilis Teodoridis:
Ulmaceae | Celtis
7. Celtis tetrandra Roxburgh; Fl. Ind.; ed. 1832. 2: 63. 1832.
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Celtis fengqingensis Hu ex E. W. Ma; C. formosana Hayata; C. kunmingensis W. C. Cheng & T. Hong; C. salvatiana C. K. Schneider; C. xizangensis E. W. Ma; C. yunnanensis C. K. Schneider.
Trees; to 30 m tall; deciduous or sometimes evergreen. Bark grayish white. Branchlets densely yellowish brown pubescent when young; brown; usually glabrescent with age; lenticels few or none. Winter buds brown; 1-3 mm; scales glabrous; margin ciliate. Stipules narrowly lanceolate; caducous; to 7 mm; never enclosing buds. Petiole 6-13 mm; brown; broadly sulcate; puberulous at least near adaxial furrow; leaf blade ovate-elliptic; ovate-lanceolate; or ± rhombic; 5-13 x 2.5-5.5 cm; thickly papery to ± papery; abaxially usually inconspicuously yellowish brown puberulent when young; with hairs scattered on major veins and sometimes tufted in vein axils with age; base oblique with one side ± rounded and other cuneate; margin entire or obtusely serrate; teeth 0-13 on each side; apex acuminate to shortly caudate-acuminate; secondary veins 3 or 4 on each side of midvein. Flowers fascicled; occasionally in racemes of 2 or 3 flowers. Style branches linear; undivided. Infructescences unbranched or seldom forked; rarely showing a small scar from a fallen male flower; 1-3 per leaf axil; rather slender; glabrous or pubescent; 0.8-1.5 cm; fruiting pedicel 1 or 2 x as long as subtending petiole. Drupe yellow to orange when mature; ± globose; 7-8 mm in diam. Stone ± globose; ca. 5 mm in diam.; 4-ribbed. Fl. Mar-Apr; fr. Sep-Oct.
Mesophytic mixed forests; valleys; slopes; 700-1500 m. W Guangxi; Hainan; Taiwan; Sichuan (Xichang); S Xizang; C; E; and S Yunnan [Bhutan; Bangladesh; India; Indonesia; Myanmar; Nepal; Thailand; Vietnam].