Vasilis Teodoridis:
Fagaceae | Lithocarpus
16. Lithocarpus handelianus A. Camus; Bull. Mus. Natl. Hist. Nat.; sér. 2. 6: 93. 1934.
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Trees to 28 m tall. Branchlets of current year stout; conspicuously sulcate; grayish brown pubescent; branchlets of previous year with yellowish brown; thick; waxy scalelike trichomes. Leaves congested at apex of branchlets; petiole 2-3 cm; sturdy with yellowish brown; thick; waxy scalelike trichomes; leaf blade elliptic to rarely ovate; 15-20 x 6-9 cm; ca. 45 x 17 cm on stump sprouts; thickly leathery; abaxially pubescent when young; with yellowish brown; thick; waxy scalelike trichomes; and pale yellowish gray; usually oily; and shiny when dry; adaxially pubescent on midvein or glabrous; base broadly cuneate to acute; margin entire; apex acute to acuminate; secondary veins 12-19 on each side of midvein; tertiary veins abaxially subparallel. Male inflorescences terminal or rarely axillary; solitary; ca. 20 cm; rachis covered with appressed short hairs. Female inflorescence with cupules in clusters of ca. 3. Infructescence rachis stout. Cupule subglobose; 2-3 cm in diam.; completely enclosing nut; apex narrowed; wall 0.5-1 mm thick; bracts imbricate; triangular-subulate; 2-4 mm; thickened; apex ± curved. Nut conical; 1.6-1.7 cm in diam.; with tawny appressed minute hairs; apex narrowed; scar ca. 1.2 cm in diam.; covering base of nut; ± concave at margin but conspicuously convex at center. Fl. May and Aug-Oct; fr. summer-autumn of following year.
* Moist sites of broad-leaved evergreen forests; 400-1000 m. Hainan