F106: Picard beech forests (Fagus sylvatica) with Hyacinthoides non-scripta, Primula acaulis, Conopodium majus

CountryFrance
Province Picardy, Normandy
References

Durin, Géhu, Noirfalise & Sougnez 1967; Lemée 1937; Roisin 1967

Remarks

CONIF
BLD
BLE
SCL
LEG
ZONPALM
ARBFERN
D-HERB
M-HERB
AZONAL WOODY
AZONAL NON-WOODY
AQUATIC
PROBLEMATIC TAXA
Sum of taxa
CONIF
0.00
BLD
9.90
BLE
0.00
SCL
2.10
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
2.60
M-HERB
10.50
AZONAL WOODY
1.00
AZONAL NON-WOODY
0.90
AQUATIC
0.00
PROBLEMATIC TAXA
1.00
Percentage of zonal taxa
CONIF
0.00
BLD
39.44
BLE
0.00
SCL
8.37
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
10.36
M-HERB
41.83
Percentage of zonal woody angiosperms
BLD
82.50
BLE
0.00
SCL
17.50
LEG
0.00
ZONPALM
0.00
Sum of all taxa28.00
Sum of zonal taxa25.10
Sum of zonal woody angiosperms12.00
Sum of % SCL + LEG17.50
Sum of % D-HERB + M-HERB (ZONAL HERB)52.19

Vegetation types

No vegetation types.

EuroVegMap information

Title

Picard beech forests (Fagus sylvatica) with Hyacinthoides non-scripta, Primula acaulis, Conopodium majus

Geographical distribution

Country/territory
France: Picardy, Normandy
area covered
33410 km²
area frequency
7 polygons .

Scientific names of main plant communities and their most common synonyms (with author citation)

Endymio-Fagetum Noirfalise & Sougnez 1967, Melico-Fagetum atlanticum Roisin 1967.

Structural feature of the main community(ies) (layers, life-forms, etc.)

In the tree layer (30 to 35 m high) there is dominant beech with admixtures of sessile oak. Their relative proportions vary quite strongly from area to area and frequently even within the same forest. pedunculate oak can also participate in the tree layer. Ash and bird cherry trees are rarer and occur in special edaphic forms in particular. A shrub layer is absent or sparse. The herb layer is species-rich and has a high degree of coverage.

Dominant and most frequent species in different layers

Tree layer
Fagus sylvatica, Quercus petraea, Fraxinus excelsior, Prunus avium
Shrub layer
often missing or only sparsely occuring: Acer pseudoplatanus, Acer campestre, Corylus avellana, Ilex aquifolium, Cornus sanguinea, Prunus avium, Crataegus laevigata, Mespilus germanica
Lianas
Lonicera periclymenum, Rubus fruticosus agg.
Herb layer
Hyacinthoides non-scripta, Galium odoratum, Melica uniflora, Euphorbia amygdaloides, Anemone nemorosa, Viola reichenbachiana, Lamium galeobdolon, Poa nemoralis, Circaea lutetiana, Milium effusum, Polygonatum multiflorum, Stellaria holostea, more rarely Neottia nidus-avis, Sanicula europaea, Vicia sepium
Moss layer (incl. lichens)
No details

Diagnostically important species

Fagus sylvatica, Quercus petraea; Hyacinthoides non-scripta, Primula acaulis, Conopodium majus (the two species mentioned last appear less frequent), Galium odoratum, Melica uniflora, Lamium galeobdolon

Ecological variants

edaphic forms:- mesotrophic form on plains or gentle slopes with a loamy covering of approximately a metre or more. It lies either on glacial drift marl, above chalk layers, or above argilliferous sands or marls; para-brown earths (luvisols) pH 5-6.- hygro-mesotrophic form on heterogeneous loams which feature a covering of loamy or marly substrates. Hygrophilous species (Circaea lutetiana, Veronica montana, Carex remota, Athyrium filix-femina, Dryopteris carthusiana and Deschampsia cespitosa) act as differential species. The B horizon is slightly gleyed and the pH lies between 5 and 5.5.- eutrophic form on deep loams that directly cover the chalky substrate or coarse lutetian limestones in the Ile de France. The profile may still number amongst the luvisol types, but its pH is between 6 to 7. This form is differentiated by Arum maculatum, Adoxa moschatellina, Ranunculus auricomus, Mercurialis perennis, Tamus communis and amongst the shrubs by Acer campestre, Euonymus europaea, Fraxinus excelsior, Ulmus minor and in the environs of Paris by Sorbus torminalis, Ligustrum vulgare, Viburnum lantana and Daphne mezereum.- acidophilous form, in which the two oaks are statistically more frequent, and which is differentiated by Holcus mollis, Convallaria majalis, Teucrium scorodonia, Polytrichum formosum and other transgressive species of the Quercion. It characterises loams with a heterogeneous covering above silicate substrates. It can be found for example in Armorican Normandy (Forest of Cérisy in the Orne), in the forest of Lyons, at the Vexin (Normandy), the Somme or the Boulonnais. Roisin (1967) raised this form to its own community (Periclymeno-Fagetum) which according to Lemée (1937) was described under the name of the Ouerceto-Fagetum using the Forest of Cérisy as an example. Luvisols with a pH of 4.5 to 5.- Form on slopes on strips of thin loam deposits that are either neutral and overgrown with Allium ursinum or acidic and overgrown with ferns such as Dryopteris filix-mas, D. carthusiana, D. dilatata, D. affinis subsp. borreri as well as Polystichum setiferum.

Geographical variants (geogr. differential species)

The Endymio-Fagetum extends from the Boulonnais up to the chalk regions of Normandy as well as via the Ile de France (Forests of Compiègne, Vellers-Cotterets etc.). In the northern range of the community (Boulonnais and Upper Normandy) two other epibiontic characteristic species appear (and in fact those of the very rare Hypericum androsaemum) as well as a species that occurs in coastal beech forests, i.e. Cardamine bulbifera (Bois de Tournehem in the Pas-de-Calais, as well as in the Bois de Lyons – Eure - to the east of Rouen). Ruscus aculeatus also appears regularly in the beech forests of Eu, Eawy and Lyons. Within the south of its range, i.e. in the south of the Seine (lower Normandy and Perche) two thermophilous species often appear on relatively dry soils: Ruscus aculeatus and Festuca heterophylla. Endymion on the other hand is restricted to fresher soils.Finally, the proportion of thermophilous species is greater in the Ile-de-France and in the Beauvaisis: Ruscus aculeatus, Festuca heterophylla, Convallaria majalis, Luzula forsteri, Sorbus torminalis. Sometimes Hypericum androsaemum is also found there.Narcissus pseudonarcissus, that appears together with Primula acaulis and Conopodium majus in the Endymio-Carpinetum, is no longer found here.

Natural accompanying vegetation (most important units in complex with the name giving mapping unit, scientific names)

The following accompanying climax communities appear in valleys which cut into chalk or lutetian limestone layers: beech forests with Fraxinus excelsior, Quercus robur and Quercus petraea (Daphno-Fagetum). These beech forests replace the communities of the Quercion pubescentis in the drier zones of the Endymio-Carpinetum above solid chalk or limestone strata. Amongst the woody plants include Ligustrum vulgare, Cornus mas, Viburnum lantana, Sorbus torminalis, Sorbus aria, Ruscus aculeatus, Daphne laureola and sometimes Buxus sempervirens. In the herb layer elements of the Cephalanthero-Fagion (Carex flacca, Carex digitata, Primula veris, Orchis mascula, Orchis purpurea, Cephalanthera damasonium, Melittis melissophyllum, Helleborus foetidus) as well as the Endymio-Fagetum (Asperula, Melica, Tamus, Endymion and Primula acaulis) are found. Important stands of these beech forests cover the chalk cliff walls of the lower Seine. The Mespilo-Quercetum of the Paris basin radiates as far as the sands located in the Perche; there are also dry grasslands.

Adjoining climax and permanent vegetation (with numbers of mapping units)

soil-acidic oak forests (F4), oak-hornbeam forests (F34), species-rich beech forests (F76), dune vegetation (P6), alder carrs (T1), alluvial and moist forests (U9, U24).

Land use, substitute communities

forestry (substitute communities, plantations of economic tree-species, scrub)
Scrub succession stages with Prunus mahaleb in particular.
grassland (meadows, pastures, herb-rich communities)
Pastures; meadows; semi dry grasslands on chalk slopes or lutetian limestones.
arable land (characteristic weed communities)
Cereal cultivation predominates in the region of Caux (Normandy), Picardy and Boulonnais, while in Lower Normandy within the bocage landscapes with low hedges, grazing and livestock breeding on pastures is much more frequent. In the region of Alençon there are high beech hedges; meadows near agricultural holdings are planted with cider apple trees.
settlements (typical ruderal vegetation)
Farms.

Site conditions of the mapping unit

Landscape type, geomorphology
Hilly country with alternations between hills and plains
Altitudinal belt and/or altitudinal range
lowland-colline; the relief does generally not excede 300 m, but in the region of Alençon in the Upper Normandie it reaches 400 m
Geology (bedrock)
limestone and siliceous rocks, till, clayey sands, chalky clay, bluff formation, flysch (Tertiary, Cretaceous)
Soil conditions
soil type, also the depth of fine textured earth (soil), stone content; (international nomenclature)
base-rich brown soils and parabrown soils; (eutric cambisols; orthic luvisols)
soil texture
loamy, clayey, partly silty, locally sandy
soil moisture, water balance
fresh to moderately dry, locally moist
soil reaction
acidic to neutral
soil fertility
mesotrophic, meso-eutrophic
Climate incl. local climate
climate type (following Walter & Lieth)
V(IV)2, VI(V)1, VI(V)2
average annual temperature
10-10,5 EC
average annual precipitation
(650)700-900 mm
average temperature of the warmest month
July 16-18 EC
average temperature of the coldest month
January 3-5 EC
local climate or other climatic peculiarities
---

Importance for nature protection

Present state and development trend of natural vegetation (degree of endangerment)
There are still well preserved semi-natural stands in state forests.
endangered communities, species worthy of protection in natural vegetation
No details
causes of endangerment
No details
required measures for protection and restitution
No details

Localities of representative stands (loci typici) with state of protection

of the natural vegetation
France: The unit exhibits very beautiful state forests, which do earn to be protected as high forests: forests of Lyons, Eu, Eawy, Hallate, Coucy, Compiègne and Viellers-Colterets.
of characteristic substitute communities worthy of protection
---

Most important references (author, year)

Durin, Géhu, Noirfalise & Sougnez 1967; Lemée 1937; Roisin 1967

Compiled by

A. Noirfalise

Taxons

Taxon
CONIF
BLD
BLE
SCL
LEG
ZONPALM
ARBFERN
D-HERB
M-HERB
AZONAL WOODY
AZONAL NON-WOODY
AQUATIC
PROBLEMATIC TAXA
CONIF
0.00
BLD
0.80
BLE
0.00
SCL
0.20
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.80
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.20
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.70
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.30
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
1.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.90
BLE
0.00
SCL
0.10
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.50
BLE
0.00
SCL
0.30
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.20
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
1.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.80
M-HERB
0.20
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
1.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.70
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.30
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.50
M-HERB
0.50
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.30
M-HERB
0.70
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
1.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
1.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
1.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
1.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.50
BLE
0.00
SCL
0.50
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.30
M-HERB
0.70
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
1.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.60
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.40
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
1.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.70
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.30
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
1.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
1.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.80
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.20
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.20
M-HERB
0.80
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.50
M-HERB
0.50
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
1.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00