F166: Oriental beech forests (Fagus sylvatica subsp. orientalis) of Talysh mountains with Hedera pastuchowii, Ilex hyrcana, Danae racemosa, alternating with deciduous broad-leaved forests (Quercus castaneifolia, Acer velutinum, Alnus subcordata)

CountryAzerbaijan
Province Talysch-mountains
References

Grossgejm 1926; Grossgejm 1948; Lebedev 1941; Prilipko 1954

Remarks

CONIF
BLD
BLE
SCL
LEG
ZONPALM
ARBFERN
D-HERB
M-HERB
AZONAL WOODY
AZONAL NON-WOODY
AQUATIC
PROBLEMATIC TAXA
Sum of taxa
CONIF
1.00
BLD
5.90
BLE
3.80
SCL
0.30
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
3.00
M-HERB
8.30
AZONAL WOODY
1.00
AZONAL NON-WOODY
0.70
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
Percentage of zonal taxa
CONIF
4.48
BLD
26.46
BLE
17.04
SCL
1.35
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
13.45
M-HERB
37.22
Percentage of zonal woody angiosperms
BLD
59.00
BLE
38.00
SCL
3.00
LEG
0.00
ZONPALM
0.00
Sum of all taxa24.00
Sum of zonal taxa22.30
Sum of zonal woody angiosperms10.00
Sum of % SCL + LEG3.00
Sum of % D-HERB + M-HERB (ZONAL HERB)50.67

Vegetation types

No vegetation types.

EuroVegMap information

Title

Oriental beech forests (Fagus sylvatica subsp. orientalis) of Talysh mountains with Hedera pastuchowii, Ilex hyrcana, Danae racemosa, alternating with deciduous broad-leaved forests (Quercus castaneifolia, Acer velutinum, Alnus subcordata)

Geographical distribution

Country/territory
Azerbaijan: Talysch-mountains
area covered
490 km²
area frequency
5 polygons .

Scientific names of main plant communities and their most common synonyms (with author citation)

No details

Structural feature of the main community(ies) (layers, life-forms, etc.)

Highly-productive, three-layered, almost pure oriental beech forests with small additions of other tree types. In the lower tree layer Taxus baccata is occasionally present. Shrub layer with Hyrcanian species, and the herb layer is herb-, grass- and fern-rich.

Dominant and most frequent species in different layers

Tree layer
Fagus sylvatica subsp. orientalis (as high as 1300 m ü. NN), with different admixture of Carpinus betulus, Tilia begoniifolia, Acer cappadocicum, Acer velutinum, Taxus baccata, Quercus castaneifolia
Shrub layer
Danae racemosa, Ilex hyrcana, Buxus hyrcana, Rubus hirtus
Lianas
Hedera pastuchowii
Herb layer
Festuca drymeja, Galium odoratum, Dryopteris filix-mas, Brachypodium sylvaticum, rare regional-specific species: Dryopteris raddeana, Scilla caucasica, Primula heterochroma, Paeonia mlokosewitschii, Polygonatum orientale, on moist sites Ranunculus villosus, on rocks Silene schafta, Sedum lencoranicum
Moss layer (incl. lichens)
No details

Diagnostically important species

Fagus sylvatica subsp. orientalis, Acer velutinum, Quercus castaneifolia, Danae racemosa, Ilex hyrcana, Buxus hyrcana, Frangula grandiflora; Dryopteris raddeana, Primula heterochroma, Paeonia mlokosewitschii, Scilla caucasica, Cephalanthera caucasica, Viola sieheana, Digitalis nervosa, further regionally specific species: Pyrus boissieriana, Pyrus grossheimii, Pyrus hyrcana, Carpinus schuschaensis

Ecological variants

Moist forms in valleys (Alnus subcordata, Pterocarya pterocarpa) and ravines.

Geographical variants (geogr. differential species)

Only altitudinal differentiation, at higher altitudes the Hyrcanian species disappear.

Natural accompanying vegetation (most important units in complex with the name giving mapping unit, scientific names)

Further down hornbeam-oriental beech forests and mixed Quercus castaneifolia forests with Acer velutinum, Parrotia persica and Zelkova carpinifolia continue on, while higher up Quercus macranthera scrub forest connects.

Adjoining climax and permanent vegetation (with numbers of mapping units)

Caucasian mixed hornbeam- oak forests (F170), Hyrcanian hygro-thermophytic mixed oak forests (H2, H3), mountain-Stipa steppes (M11), thorn-cushion vegetation (N8).

Land use, substitute communities

forestry (substitute communities, plantations of economic tree-species, scrub)
Forestry
grassland (meadows, pastures, herb-rich communities)
Pastures.
arable land (characteristic weed communities)
---
settlements (typical ruderal vegetation)
---

Site conditions of the mapping unit

Landscape type, geomorphology
Very varied mountainous relief: north exposed mountain slopes and shady ravines
Altitudinal belt and/or altitudinal range
montane to altimontane; (600) 900-1800 m
Geology (bedrock)
sediment and eruptive rocks (Tertiary)
Soil conditions
soil type, also the depth of fine textured earth (soil), stone content; (international nomenclature)
different formations of brown mountain-forest soils; (eutric cambisols; locally leptosols)
soil texture
loamy, sandy-loamy, mostly stone-rich
soil moisture, water balance
fresh to moist
soil reaction
moderately acidic to neutral
soil fertility
meso-eutrophic
Climate incl. local climate
climate type (following Walter & Lieth)
X
average annual temperature
6-12 EC
average annual precipitation
(650) 800-1300 mm
average temperature of the warmest month
18-22 EC
average temperature of the coldest month
-5 to -1,2 EC
local climate or other climatic peculiarities
Unevenly distributed yearly precipitation: moist autumn and winter, dry and warm summer. The dryness of the climate grows with the rising sea level.

Importance for nature protection

Present state and development trend of natural vegetation (degree of endangerment)
Northern outliers of the widespread Hyrcanian beech forests that have their distribution strongholds in the Elburs mountains of northern Iran. Forest stands damaged by timber harvesting and forest grazing; they have important soil and water protective functions, and are of great importance for nature conservation (relict forests, numerous endemics).
endangered communities, species worthy of protection in natural vegetation
Endemic species and relict occurences on the Talysch-mountains: Ilex hyrcana, Buxus hycana, Taxus baccata.
causes of endangerment
Clearing, timber harvesting, forest grazing.
required measures for protection and restitution
The remnants of the Hyrcanian beech forests should be safeguarded in a large "Hyrcanian nature reserve".

Localities of representative stands (loci typici) with state of protection

of the natural vegetation
Azerbaijan: nature reserve Girkan.
of characteristic substitute communities worthy of protection
---

Most important references (author, year)

Grossgejm 1926; Grossgejm 1948; Lebedev 1941; Prilipko 1954

Compiled by

Z. Neuhäuslová based on documents by A.G. Doluchanov and Ju.R. Šeljag-Sosonko

Taxons

Taxon
CONIF
BLD
BLE
SCL
LEG
ZONPALM
ARBFERN
D-HERB
M-HERB
AZONAL WOODY
AZONAL NON-WOODY
AQUATIC
PROBLEMATIC TAXA
CONIF
0.00
BLD
0.80
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.20
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.90
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.10
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.80
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.20
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
1.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
1.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
1.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
1.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
1.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
1.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
1.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.50
M-HERB
0.50
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.80
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.20
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
1.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.10
M-HERB
0.60
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.30
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.40
M-HERB
0.60
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.20
M-HERB
0.80
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
1.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.70
M-HERB
0.30
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.50
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.50
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.80
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.20
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.80
M-HERB
0.20
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.30
M-HERB
0.70
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
1.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.70
BLE
0.00
SCL
0.30
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00