G56: Balkanic-Rhodopean Oriental hornbeam-hop-hornbeam forests (Ostrya carpinifolia, Carpinus orientalis), partly with Haberlea rhodopensis

CountryBulgaria, Greece
Province Bulgaria
References

Bondev 1991

Remarks

CONIF
BLD
BLE
SCL
LEG
ZONPALM
ARBFERN
D-HERB
M-HERB
AZONAL WOODY
AZONAL NON-WOODY
AQUATIC
PROBLEMATIC TAXA
Sum of taxa
CONIF
4.00
BLD
9.80
BLE
0.50
SCL
1.70
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
3.50
M-HERB
3.30
AZONAL WOODY
2.00
AZONAL NON-WOODY
0.20
AQUATIC
0.00
PROBLEMATIC TAXA
2.00
Percentage of zonal taxa
CONIF
17.54
BLD
42.98
BLE
2.19
SCL
7.46
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
15.35
M-HERB
14.47
Percentage of zonal woody angiosperms
BLD
81.67
BLE
4.17
SCL
14.17
LEG
0.00
ZONPALM
0.00
Sum of all taxa27.00
Sum of zonal taxa22.80
Sum of zonal woody angiosperms12.00
Sum of % SCL + LEG14.17
Sum of % D-HERB + M-HERB (ZONAL HERB)29.82

Vegetation types

No vegetation types.

EuroVegMap information

Title

Balkanic-Rhodopean Oriental hornbeam-hop-hornbeam forests (Ostrya carpinifolia, Carpinus orientalis), partly with Haberlea rhodopensis

Geographical distribution

Country/territory
Small-scaled: Bulgaria: north Rhodopes, Balkan-mountains; north-eastern Greece: Nestos-valley
area covered
140 km²
area frequency
2 polygons.

Scientific names of main plant communities and their most common synonyms (with author citation)

Ostryeta carpinifoliae Bondev 1991, Ostryo-Carpinion Horvat 1959.

Structural feature of the main community(ies) (layers, life-forms, etc.)

Low-growing, bush-formed, usually two-layered (tree and herb layer) forests with many thermophilous species.

Dominant and most frequent species in different layers

Tree layer
Ostrya carpinifolia, Carpinus orientalis, Fraxinus ornus, Quercus pubescens, Acer campestre, Fraxinus excelsior, (Pinus nigra, Pinus sylvestris, Fagus sylvatica subsp. moesiaca, Populus tremula)
Shrub layer
Crataegus monogyna, Rosa pimpinellifolia, Juniperus communis, Juniperus oxycedrus, Sorbus aria, Cornus mas
Lianas
Hedera helix, Clematis vitalba
Herb layer
Sesleria latifolia, Cyclamen hederifolium, Erythronium dens-canis, Geranium sanguineum, Clinopodium vulgare, Brachypodium sylvaticum, Trifolium alpestre, Geum urbanum, Festuca pirinensis
Moss layer (incl. lichens)
Hypnum cupressiforme, Hylocomium splendens

Diagnostically important species

Ostrya carpinifolia, Carpinus orientalis, Quercus pubescens; Sesleria latifolia, Festuca pirinensis, Haberlea rhodopensis, Erythronium dens-canis

Ecological variants

---

Geographical variants (geogr. differential species)

Only in the Rhodope mountains with Haberlea rhodopensis.

Natural accompanying vegetation (most important units in complex with the name giving mapping unit, scientific names)

Forests with Fagus sylvatica subsp. moesiaca, Quercus dalechampii, Pinus nigra, Quercus cerris; alluvial forests, as well as Carpinion orientalis communities.

Adjoining climax and permanent vegetation (with numbers of mapping units)

Pine forests (D63), species-rich beech forests (F153, F154), sessile oak forests (G9), Balkan oak forests (G28), hop-hornbeam forests (G56), Oriental hornbeam-downy oak forests (G59), black pine forests (K16), alluvial forests (U18).

Land use, substitute communities

forestry (substitute communities, plantations of economic tree-species, scrub)
Degraded shrub forests with Carpinus orientalis, Ostrya carpinifolia, Juniper species, Pinus forestry.
grassland (meadows, pastures, herb-rich communities)
Sub-Mediterranean dry grasslands.
arable land (characteristic weed communities)
---
settlements (typical ruderal vegetation)
---

Site conditions of the mapping unit

Landscape type, geomorphology
Sunny ravine slopes with rocky projections and slopes of river valleys where higher humidity prevails.
Altitudinal belt and/or altitudinal range
colline to montane; 100-1300 m
Geology (bedrock)
carbonate rocks (Cretaceous), siliceous rock (Precambrian)
Soil conditions
soil type, also the depth of fine textured earth (soil), stone content; (international nomenclature)
cinnamonic and grey forest soils; humus carbonate soils with high proportion of raw stone content; (chromic and humic cambisols; chromic luvisols; rendzic leptosols)
soil texture
loamy, stony
soil moisture, water balance
fresh to dry
soil reaction
acidic to neutral
soil fertility
oligo-mesotrophic, mesotrophic
Climate incl. local climate
climate type (following Walter & Lieth)
VI(VII), X
average annual temperature
9-11 °C
average annual precipitation
700 mm
average temperature of the warmest month
July 18-22 °C
average temperature of the coldest month
January -2 to 0 °C
local climate or other climatic peculiarities
Strong insolation on south-exposed slopes, high air humidity in gorges and on valley slopes.

Importance for nature protection

Present state and development trend of natural vegetation (degree of endangerment)
Quite well preserved, not used for forestry.
endangered communities, species worthy of protection in natural vegetation
All stands are worthy of protection.
causes of endangerment
---
required measures for protection and restitution
Many stands protected in the Rhodope mountains and the Balkan mountains.

Localities of representative stands (loci typici) with state of protection

of the natural vegetation
Bulgaria: Červenata stena, Džendema, Soteneto; Greece: Nestos-gorge near Toxates, Nestos-upper course at the Bulgarian border.
of characteristic substitute communities worthy of protection
---

Most important references (author, year)

Bondev 1991

Compiled by

N. Doniţă based on documents by I. Bondev and Z. Neuhäuslová, Th. Raus and E. Bergmeier

Taxons

Taxon
CONIF
BLD
BLE
SCL
LEG
ZONPALM
ARBFERN
D-HERB
M-HERB
AZONAL WOODY
AZONAL NON-WOODY
AQUATIC
PROBLEMATIC TAXA
CONIF
0.00
BLD
0.80
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.20
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.50
BLE
0.00
SCL
0.50
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.50
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.50
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.50
BLE
0.00
SCL
0.50
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
1.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.20
M-HERB
0.80
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.60
M-HERB
0.40
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
1.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.40
M-HERB
0.60
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.70
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.30
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.80
BLE
0.00
SCL
0.20
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.70
M-HERB
0.30
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.20
M-HERB
0.60
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.20
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.50
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.50
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
1.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
1.00
CONIF
1.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
1.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
1.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
1.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
1.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.50
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.50
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.80
BLE
0.00
SCL
0.20
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.70
BLE
0.00
SCL
0.30
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.90
M-HERB
0.10
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
1.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.50
M-HERB
0.50
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00