M6: Central Pontic herb-grass steppes (Stipa lessingiana, Stipa capillata, Salvia nutans, Salvia tesquicola, Medicago falcata subsp. romanica) with Calophaca wolgarica, Stipa zalesskii, Salvia stepposa, Aster sedifolius subsp. angustissimus

CountryRussian Federation, Ukraine
Province Don- and Choper-river floodplains and southern slopes of the central Russian and Volgian highlands. Within the broad strip along the lower courses of the Severskij Donec, Derkul, Kalitva and Bystraja in the west to the upper course of the Medvedica in the north-east and the Volga in the east
References

Bilyk 1973b; Bilyk & Tkačenko 1971; Dobročaeva 1956; Keller (Red.) 1931; Lavrenko 1940; Lavrenko 1980; Lavrenko & Dochman 1933; Tarasov 1958; Tarasov 1968; Tkačenko 1989

Remarks

CONIF
BLD
BLE
SCL
LEG
ZONPALM
ARBFERN
D-HERB
M-HERB
AZONAL WOODY
AZONAL NON-WOODY
AQUATIC
PROBLEMATIC TAXA
Sum of taxa
CONIF
0.00
BLD
2.00
BLE
0.00
SCL
0.50
LEG
0.50
ZONPALM
0.00
ARBFERN
0.00
D-HERB
7.30
M-HERB
3.70
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
Percentage of zonal taxa
CONIF
0.00
BLD
14.29
BLE
0.00
SCL
3.57
LEG
3.57
ZONPALM
0.00
ARBFERN
0.00
D-HERB
52.14
M-HERB
26.43
Percentage of zonal woody angiosperms
BLD
66.67
BLE
0.00
SCL
16.67
LEG
16.67
ZONPALM
0.00
Sum of all taxa14.00
Sum of zonal taxa14.00
Sum of zonal woody angiosperms3.00
Sum of % SCL + LEG33.33
Sum of % D-HERB + M-HERB (ZONAL HERB)78.57

Vegetation types

No vegetation types.

EuroVegMap information

Title

Central Pontic herb-grass steppes (Stipa lessingiana, Stipa capillata, Salvia nutans, Salvia tesquicola, Medicago falcata subsp. romanica) with Calophaca wolgarica, Stipa zalesskii, Salvia stepposa, Aster sedifolius subsp. angustissimus

Geographical distribution

Country/territory
Ukraine; Russia: Don- and Choper-river floodplains and southern slopes of the central Russian and Volgian highlands. Within the broad strip along the lower courses of the Severskij Donec, Derkul, Kalitva and Bystraja in the west to the upper course of the Medvedica in the north-east and the Volga in the east
area covered
55550 km²
area frequency
14 polygons.

Scientific names of main plant communities and their most common synonyms (with author citation)

Stipeta lessingianae herbosa, Stipeta capillata herbosa Lavrenko 1940.

Structural feature of the main community(ies) (layers, life-forms, etc.)

The dominant community types are herb-rich Festuca valesiaca-Stipa steppes with Pontic-Kazakh and western paleoarctic xerophytic tussock grasses (Stipa lessingiana, Stipa capillata, Festuca valesiaca) and with the east Pontic-Kazakh species Stipa zalesskii. Widespread species include Bromus riparius, Koeleria macrantha, Agropyron cristatum subsp. pectinatum, Elymus hispidus, Elymus hispidus subsp. barbulatus, and one also finds Poa angustifolia occasionally, and Poa bulbosa at impeded sites. Stipa pulcherrima, Stipa tirsa, and Stipa dasyphylla are much rarer. The absence of Stipa ucrainica, that dominates in the west-Pontic steppes, is characteristic. Xerophytic steppe species dominate amongst the herbs. Many "eastern" elements (mainly east-Pontic-Kazakh species) are typical here: Dianthus leptopetalus, Onosma simplicissima, Salvia stepposa, Aster sedifolius subsp. angustissimus, Aster tarbagatensis, Tanacetum achilleifolium, Onosma arenaria etc. Ephemerals and ephemeroids. Shrubs are also always present.

Dominant and most frequent species in different layers

Tree layer
---
Shrub layer
Caragana frutex, Calophaca wolgarica, Prunus tenella
Lianas
---
Herb layer
Stipa lessingiana, Stipa capillata, Festuca valesiaca, Agropyron cristatum subsp. pectinatum, Elymus hispidus, Serratula erucifolia, Seseli campestre, Dianthus leptopetalus, Holosteum umbellatum, Gagea pusilla, Poa bulbosa
Moss layer (incl. lichens)
No details

Diagnostically important species

Stipa lessingiana, Stipa zalesskii, Calophaca wolgarica, Salvia stepposa, Salvia tesquicola, Medicago falcata subsp. romanica

Ecological variants

---

Geographical variants (geogr. differential species)

In the east of the range Calophaca wolgarica; on solonetz soils and solonetz-like black earths Artemisia lerchiana and Artemisia pauciflora.

Natural accompanying vegetation (most important units in complex with the name giving mapping unit, scientific names)

Small-area incidences of petrophytic herb grass steppes (Stipa tirsa, Stipa zalesskii, Stipa dasyphylla, Stipa pulcherrima, Festuca valesiaca, Achillea nobilis, Teucrium capitatum, Ephedra distachya) on slate and sandstone in the Donets area; tomillares and tomillares steppes (Koeleria talievii, Stipa cretacea, Erysimum cretaceum, Daphne sophia, Thymus calcareus, Hyssopus cretaceus, Artemisia hololeuca) on chalk and marlstone in the area of the Donets and Don rivers; east European open pedunculate oak forests (Quercus robur) with steppe species (Caragana frutex, Prunus tenella, Brachypodium pinnatum) at valley slopes; Festuca valesiaca-steppes in complex with halophytic dwarf semi-shrubs.

Adjoining climax and permanent vegetation (with numbers of mapping units)

pine forests (D57), lime-pedunculate oak forests (F72), forest steppes (L5), steppes (M1, M2, M5, M7, M8, M9, M13).

Land use, substitute communities

forestry (substitute communities, plantations of economic tree-species, scrub)
---
grassland (meadows, pastures, herb-rich communities)
Pastures, halophilous vegetation.
arable land (characteristic weed communities)
Wheat fields.
settlements (typical ruderal vegetation)
---

Site conditions of the mapping unit

Landscape type, geomorphology
river floodplains with a primarily flat relief; slightly undulating highlands with ravines and river valleys, slopes
Altitudinal belt and/or altitudinal range
lowland; 100-150 m (max. 220-230 m)
Geology (bedrock)
eolian (sand, bluff formation) and glacial sediments (Pleistocene above Tertiary, Cretaceous)
Soil conditions
soil type, also the depth of fine textured earth (soil), stone content; (international nomenclature)
few humic black earths; (haplic and luvic sodic chernozems; haplic sodic kastanozems)
soil texture
sandy-loamy, loamy-sandy
soil moisture, water balance
moderately dry
soil reaction
neutral
soil fertility
meso-eutrophic
Climate incl. local climate
climate type (following Walter & Lieth)
VII1a
average annual temperature
6-8 °C
average annual precipitation
350-470 mm
average temperature of the warmest month
22-25 °C
average temperature of the coldest month
-10 to -7 °C
local climate or other climatic peculiarities
continental climate with frequent droughts, dry winds, hot dry summers and cool winters.

Importance for nature protection

Present state and development trend of natural vegetation (degree of endangerment)
At interfluvial sites these steppes have been transformed largely into arable land. Their remnants are associated with sites of little agricultural use. With absolute protective regimes the grass steppe is transformed into a shrub steppe with dominant Caragana frutex.
endangered communities, species worthy of protection in natural vegetation
Worthy of protection and highly valuable are above all tomillares on Cretaceous-disclosures at steep slopes of the river valleys.
causes of endangerment
Agricultural use.
required measures for protection and restitution
Establishment of suitable management plans for protected areas.

Localities of representative stands (loci typici) with state of protection

of the natural vegetation
Ukraine: Strel'covskaja steppe in the Lugansk-region.
of characteristic substitute communities worthy of protection
---

Most important references (author, year)

Bilyk 1973b; Bilyk & Tkačenko 1971; Dobročaeva 1956; Keller (Red.) 1931; Lavrenko 1940; Lavrenko 1980; Lavrenko & Dochman 1933; Tarasov 1958; Tarasov 1968; Tkačenko 1989

Compiled by

Z. Neuhäuslová based on documents by Z.V. Karamyševa

Taxons

Taxon
CONIF
BLD
BLE
SCL
LEG
ZONPALM
ARBFERN
D-HERB
M-HERB
AZONAL WOODY
AZONAL NON-WOODY
AQUATIC
PROBLEMATIC TAXA
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.80
M-HERB
0.20
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.50
BLE
0.00
SCL
0.00
LEG
0.50
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.50
BLE
0.00
SCL
0.50
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.60
M-HERB
0.40
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.70
M-HERB
0.30
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.40
M-HERB
0.60
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.70
M-HERB
0.30
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.60
M-HERB
0.40
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.50
M-HERB
0.50
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
1.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.60
M-HERB
0.40
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.80
M-HERB
0.20
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.70
M-HERB
0.30
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.90
M-HERB
0.10
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00