M8: Pontic psammophytic herb-grass steppes (Stipa anomala, Agropyron spp., Festuca beckeri, Koeleria sabuletorum, Leymus racemosus, Helichrysum arenarium, Euphorbia seguieriana) with Artemisia marschalliana

CountryRussian Federation, Ukraine
Province river carved valleys, especially of Dnjepr and Don, along the middle and lower courses of the Dnjepr (Kachovskaja, Kozačelagerskaja, Vinogradovsaja, Zbur'ecskaja, Ivanovskaja areny and Kinburnskaja-bay-mouth bar). Along the Don there are large sand massifs (near Cimljansk (partly flooded) and Arčedinskie peski)
References

Bilyk 1956; Gael’ 1932; Gordienko 1969; Gožev 1929; Kleopov 1934; Konstantinova 1963; Kosec’ & Tkačenko 1973; Lavrenko 1927; Lavrenko 1935; Lavrenko & Poreckij 1928; Pačoskij 1923b; Vicherek 1972

Remarks

CONIF
BLD
BLE
SCL
LEG
ZONPALM
ARBFERN
D-HERB
M-HERB
AZONAL WOODY
AZONAL NON-WOODY
AQUATIC
PROBLEMATIC TAXA
Sum of taxa
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
9.60
M-HERB
2.20
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.20
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
Percentage of zonal taxa
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
81.36
M-HERB
18.64
Percentage of zonal woody angiosperms
BLD
NaN
BLE
NaN
SCL
NaN
LEG
NaN
ZONPALM
NaN
Sum of all taxa12.00
Sum of zonal taxa11.80
Sum of zonal woody angiosperms0.00
Sum of % SCL + LEGNaN
Sum of % D-HERB + M-HERB (ZONAL HERB)100.00

Vegetation types

No vegetation types.

EuroVegMap information

Title

Pontic psammophytic herb-grass steppes (Stipa anomala, Agropyron spp., Festuca beckeri, Koeleria sabuletorum, Leymus racemosus, Helichrysum arenarium, Euphorbia seguieriana) with Artemisia marschalliana

Geographical distribution

Country/territory
Ukraine, Russia: river carved valleys, especially of Dnjepr and Don, along the middle and lower courses of the Dnjepr (Kachovskaja, Kozačelagerskaja, Vinogradovsaja, Zbur'ecskaja, Ivanovskaja areny and Kinburnskaja-bay-mouth bar). Along the Don there are large sand massifs (near Cimljansk (partly flooded) and Arčedinskie peski)
area covered
6570 km²
area frequency
30 polygons.

Scientific names of main plant communities and their most common synonyms (with author citation)

Festucion beckeri Vicherek 1972; Stipeta borysthenicae arenoso-herbosa.

Structural feature of the main community(ies) (layers, life-forms, etc.)

Psammophytic steppe grasses (Stipa anomala, Festuca beckeri, rarely Koeleria sabuletorum) as well as psammophytic herbs with dominating and the most important stand forming species (e.g. Euphorbia seguieriana, Thymus kosteleckyanus, Helichrysum arenarium, Dianthus platyodon, Goniolimon graminifolium, Asperula graveolens, Achillea micrantha, Scorzonera ensifolia, Jurinea longifolia). Other obligatory psammophytes and annuals are frequently present.

Dominant and most frequent species in different layers

Tree layer
---
Shrub layer
---
Lianas
---
Herb layer
Stipa anomala, Festuca beckeri, Koeleria sabuletorum, Goniolimon graminifolium, Asperula graveolens, Achillea micrantha, Scorzonera ensifolia, Silene borysthenica, Syrenia montanea, Onosma arenaria, Plantago arenaria, Polygonum arenarium
Moss layer (incl. lichens)
No details

Diagnostically important species

Stipa anomala, Festuca beckeri, Koeleria sabuletorum, Leymus racemosus, Helichrysum arenarium, Euphorbia seguieriana, Artemisia marschalliana

Ecological variants

---

Geographical variants (geogr. differential species)

In the species assemblage of the Dnieper steppes one finds Agropyron dasyanthum, Thymus borysthenicus, Centaurea margeritacea subsp. breviceps, in the Don steppes Agropyron tanaiticum, Thymus pallasianus, Centaurea margeritacea subsp. protogerberi, Centaurea margeritacea subsp. dubjanskyi, Centaurea margeritacea subsp. gerberi and in the steppes near Arčedinskie peski Juniperus sabina. The sand terraces of the Dnieper are inhabited by pine forests (Pinus sylvestris).

Natural accompanying vegetation (most important units in complex with the name giving mapping unit, scientific names)

Oak (Quercus robur), birch (Betula pendula) or black alder forests, pine forests, tall reed vegetation and tall sedge communities.

Adjoining climax and permanent vegetation (with numbers of mapping units)

pine forests (D57), steppes (M1, M2, M5, M6, M7, M12, M13), desert steppes (M16), dune vegetation (P15), salt vegetation (P30, P34), freshwater-tall reed banks (R1), alluvial forests (U14).

Land use, substitute communities

forestry (substitute communities, plantations of economic tree-species, scrub)
---
grassland (meadows, pastures, herb-rich communities)
Pasture grounds.
arable land (characteristic weed communities)
Wheat fields.
settlements (typical ruderal vegetation)
---

Site conditions of the mapping unit

Landscape type, geomorphology
floodplain terraces of old alluvial sand sediments; slightly undulating hilly country
Altitudinal belt and/or altitudinal range
lowland; 0-200 m
Geology (bedrock)
eolian (sand, bluff formation) and fluviatile sediments (Pleistocene, Pleistocene above Tertiary, Cretaceous)
Soil conditions
soil type, also the depth of fine textured earth (soil), stone content; (international nomenclature)
southern black earths and dark chestnut soils; (haplic sodic kastanozems; haplic (sodic) chernozems)
soil texture
sandy
soil moisture, water balance
moderately dry
soil reaction
neutral
soil fertility
mesotrophic
Climate incl. local climate
climate type (following Walter & Lieth)
VII1a
average annual temperature
7-10 °C
average annual precipitation
300-350 mm
average temperature of the warmest month
21-22 °C
average temperature of the coldest month
-7 to -5 °C
local climate or other climatic peculiarities
moderate continental climate.

Importance for nature protection

Present state and development trend of natural vegetation (degree of endangerment)
A large part of the steppes has been converted into arable land, and many parts have been damaged severely by grazing.
endangered communities, species worthy of protection in natural vegetation
near-natural steppe communities.
causes of endangerment
Intense grazing, agricultural use.
required measures for protection and restitution
Protection from overgrazing.

Localities of representative stands (loci typici) with state of protection

of the natural vegetation
Ukraine: in two areas of the biosphere reserve and Zapovednik "Černomorskij" on the Kinburnskaja bay-mouth bar.
of characteristic substitute communities worthy of protection
---

Most important references (author, year)

Bilyk 1956; Gael’ 1932; Gordienko 1969; Gožev 1929; Kleopov 1934; Konstantinova 1963; Kosec’ & Tkačenko 1973; Lavrenko 1927; Lavrenko 1935; Lavrenko & Poreckij 1928; Pačoskij 1923b; Vicherek 1972

Compiled by

Z. Neuhäuslová based on documents by Z.V. Karamyševa

© Bundesamt für Naturschutz (BfN)
Texts were extracted from the EuroVegMap application with permission from the authors.

Taxons

Taxon
CONIF
BLD
BLE
SCL
LEG
ZONPALM
ARBFERN
D-HERB
M-HERB
AZONAL WOODY
AZONAL NON-WOODY
AQUATIC
PROBLEMATIC TAXA
Taxon
Achillea micrantha
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.70
M-HERB
0.30
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
Taxon
Asperula graveolens
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.00
M-HERB
0.80
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.20
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
Taxon
Festuca beckeri
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
1.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
Taxon
Goniolimon graminifolium
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.70
M-HERB
0.30
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
Taxon
Koeleria sabuletorum
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.80
M-HERB
0.20
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
Taxon
Onosma arenaria
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.80
M-HERB
0.20
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
Taxon
Plantago arenaria
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
1.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
Taxon
Polygonum arenarium
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
1.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
Taxon
Scorzonera ensifolia
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.90
M-HERB
0.10
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
Taxon
Silene borysthenica
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.80
M-HERB
0.20
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
Taxon
Stipa anomala
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
1.00
M-HERB
0.00
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00
Taxon
Syrenia montanea
CONIF
0.00
BLD
0.00
BLE
0.00
SCL
0.00
LEG
0.00
ZONPALM
0.00
ARBFERN
0.00
D-HERB
0.90
M-HERB
0.10
AZONAL WOODY
0.00
AZONAL NON-WOODY
0.00
AQUATIC
0.00
PROBLEMATIC TAXA
0.00

Building of the IPR-vegetation analysis website and database was supported by the grant projects GA ČR (Grant Agency of the Czech Republic) No. P210/10/0124 and 18-25057S.